1 Life Sciences Institute, University of Michigan, Ann Arbor, Michigan, United States of America, 2 Department of Cell and Developmental Biology, University of Michigan, Ann Arbor, Michigan, United States of America, 3 Department of Pharmacology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America, 4 Michigan Neuroscience Institute, University of Michigan Medical School, Ann Arbor, Michigan, United States of美国,美国密歇根大学医学院5人类遗传学系,美国密歇根州安阿伯市,美国,美国6号神经生物学系,杜克大学医学中心,北卡罗来纳州达勒姆大学医学中心,美国,美国北卡罗来纳州7号,杜克大学生物医学工程系美国,美国密歇根大学医学院精神病学系9